Brash space adventurer Peter Quill (Chris Pratt) finds himself the quarry of relentless bounty hunters after he steals an orb coveted by Ronan, a powerful villain. To evade Ronan, Quill is forced into an uneasy truce with four disparate misfits: gun-toting Rocket Raccoon, treelike-humanoid Groot, enigmatic Gamora, and vengeance-driven Drax the Destroyer. But when he discovers the orb’s true power and the cosmic threat it poses, Quill must rally his ragtag group to save the universe.
Peter Quill and his fellow Guardians are hired by a powerful alien race, the Sovereign, to protect their precious batteries from invaders. When it is discovered that Rocket has stolen the items they were sent to guard, the Sovereign dispatch their armada to search for vengeance. As the Guardians try to escape, the mystery of Peter's parentage is revealed.