Convinced that mutants pose a threat to humanity, Dr. Bolivar Trask (Peter Dinklage) develops the Sentinels, enormous robotic weapons that can detect a mutant gene and zero in on that person. In the 21st century, the Sentinels have evolved into highly efficient killing machines. With mutants now facing extinction, Wolverine (Hugh Jackman) volunteers to go back in time and rally the X-Men of the past to help change a pivotal moment in history and thereby save their future.
Peter Quill and his fellow Guardians are hired by a powerful alien race, the Sovereign, to protect their precious batteries from invaders. When it is discovered that Rocket has stolen the items they were sent to guard, the Sovereign dispatch their armada to search for vengeance. As the Guardians try to escape, the mystery of Peter's parentage is revealed.